About the Authors

Matin Miryeganeh

Affiliation Department of Biology, Graduate School of Science, Chiba University, 1-33 Yayoi, Inage, Chiba, Japan

Koji Takayama

Affiliation The University Museum, The University of Tokyo, 7-3-1 Hongo, Bunkyo-ku, Tokyo, Japan

Yoichi Tateishi

Affiliation Faculty of Education, University of the Ryukyus, 1 Senbaru, Nakagami-gun Okinawa, Japan

Tadashi Kajita

tkaji@faculty.chiba-u.jp

Affiliation Department of Biology, Graduate School of Science, Chiba University, 1-33 Yayoi, Inage, Chiba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MM TK. Performed the experiments: MM TK. Analyzed the data: MM TK. Contributed reagents/materials/analysis tools: KT YT TK. Wrote the paper: MM KT TK. Designed the whole project including this study: KT YT TK.