About the Authors

Doo-Hyung Lee

dl343@cornell.edu

Current address: Department of Life Sciences, Gachon University, Seongnam-si, Gyeonggi-do, South Korea

Affiliation U.S. Department of Agriculture – Agricultural Research Service, Appalachian Fruit Research Station, Kearneysville, West Virginia, United States of America

John P. Cullum

Affiliation Department of Entomology, Virginia Tech, Winchester, Virginia, United States of America

Jennifer L. Anderson

Affiliation U.S. Department of Agriculture – Animal and Plant Health Inspection Service, National Detector Dog Training Center, Newnan, Georgia, United States of America

Jodi L. Daugherty

Affiliation U.S. Department of Agriculture – Animal and Plant Health Inspection Service, National Detector Dog Training Center, Newnan, Georgia, United States of America

Lisa M. Beckett

Affiliation U.S. Department of Agriculture – Animal and Plant Health Inspection Service, National Detector Dog Training Center, Newnan, Georgia, United States of America

Tracy C. Leskey

Affiliation U.S. Department of Agriculture – Agricultural Research Service, Appalachian Fruit Research Station, Kearneysville, West Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: DHL TCL LMB. Performed the experiments: DHL JPC JLA JLD. Analyzed the data: DHL TCL. Contributed reagents/materials/analysis tools: TCL LMB. Wrote the paper: DHL TCL JLA JLD.