About the Authors

Barbara Dema

Affiliation Laboratory of Molecular Immunogenetics, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Christophe Pellefigues

Affiliation Institut National de la Santé et de la Recherche Médicale U699, Université Paris Diderot, Paris, France

Sarfaraz Hasni

Affiliation Office of the Clinical Director, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Nathalie Gault

Affiliation Clinical Research Unit, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Chao Jiang

Affiliation Laboratory of Molecular Immunogenetics, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Tiffany K. Ricks

Affiliation Laboratory of Molecular Immunogenetics, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Michael M. Bonelli

Affiliation Lymphocyte Biology Section, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Jörg Scheffel

Affiliation Laboratory of Molecular Immunogenetics, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Karim Sacré

Affiliations Institut National de la Santé et de la Recherche Médicale U699, Université Paris Diderot, Paris, France, Department of Internal Medicine, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Mathieu Jablonski

Affiliation Department of Nephrology, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Delphine Gobert

Affiliation Department of Internal Medicine, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Thomas Papo

Affiliations Institut National de la Santé et de la Recherche Médicale U699, Université Paris Diderot, Paris, France, Department of Internal Medicine, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Eric Daugas

Affiliations Institut National de la Santé et de la Recherche Médicale U699, Université Paris Diderot, Paris, France, Department of Nephrology, Hôpital Bichat, Assistance Publique-Hôpitaux de Paris, Université Paris Diderot, Faculté de Médecine site Bichat, Paris, France

Gabor Illei

Affiliation Sjogren’s Syndrome Clinic, National Institute of Dental and Craniofacial Research, National Institutes of Health, Bethesda, Maryland, United States of America

Nicolas Charles

Contributed equally to this work with: Nicolas Charles, Juan Rivera

juan_rivera@nih.gov (JR); nicolas.charles@inserm.fr (NC)

Affiliation Institut National de la Santé et de la Recherche Médicale U699, Université Paris Diderot, Paris, France

Juan Rivera

Contributed equally to this work with: Nicolas Charles, Juan Rivera

juan_rivera@nih.gov (JR); nicolas.charles@inserm.fr (NC)

Affiliation Laboratory of Molecular Immunogenetics, National Institute of Arthritis and Musculoskeletal and Skin Diseases, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BD NC SH ED GI JR. Performed the experiments: BD CP MMB JS CJ TKR. Analyzed the data: BD NG MMB SH GI NC JR. Contributed reagents/materials/analysis tools: SH KS MJ DG TP ED. Wrote the paper: BD SH NG MJ DG ED GI NC JR.