About the Authors

Candice C. Clay

Contributed equally to this work with: Candice C. Clay, Kinjal Maniar-Hew

Affiliation California National Primate Research Center, University of California Davis, Davis, California, United States of America

Kinjal Maniar-Hew

Contributed equally to this work with: Candice C. Clay, Kinjal Maniar-Hew

Affiliation California National Primate Research Center, University of California Davis, Davis, California, United States of America

Joan E. Gerriets

Affiliation California National Primate Research Center, University of California Davis, Davis, California, United States of America

Theodore T. Wang

Affiliation California National Primate Research Center, University of California Davis, Davis, California, United States of America

Edward M. Postlethwait

Affiliation Department of Environmental Health Sciences, School of Public Health, University of Alabama, Birmingham, Alabama, United States of America

Michael J. Evans

Affiliations California National Primate Research Center, University of California Davis, Davis, California, United States of America, Department of Anatomy, Physiology, and Cell Biology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Justin H. Fontaine

Affiliation California National Primate Research Center, University of California Davis, Davis, California, United States of America

Lisa A. Miller

lmiller@ucdavis.edu

Affiliations California National Primate Research Center, University of California Davis, Davis, California, United States of America, Department of Anatomy, Physiology, and Cell Biology, School of Veterinary Medicine, University of California Davis, Davis, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CCC KMH EMP MJE LAM. Performed the experiments: CCC KMH TTW JEG JHF. Analyzed the data: CCC KMH TTW JEG LAM. Wrote the paper: CCC KMH LAM. Assay development: JHF.