About the Authors

Shawna J. Cikanek

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Simon Nockold

Affiliation Panama Amphibian Rescue and Conservation Project, Smithsonian Tropical Research Institute, Gamboa, Republic of Panama

Janine L. Brown

Affiliation Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

James W. Carpenter

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Kansas State University, Manhattan, Kansas, United States of America

Angie Estrada

Affiliation Panama Amphibian Rescue and Conservation Project, Smithsonian Tropical Research Institute, Gamboa, Republic of Panama

Jorge Guerrel

Affiliation Panama Amphibian Rescue and Conservation Project, Smithsonian Tropical Research Institute, Gamboa, Republic of Panama

Katharine Hope

Affiliation Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Roberto Ibáñez

Affiliation Panama Amphibian Rescue and Conservation Project, Smithsonian Tropical Research Institute, Gamboa, Republic of Panama

Sarah B. Putman

Affiliation Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Brian Gratwicke

gratwickeb@si.edu

Affiliation Smithsonian Conservation Biology Institute, National Zoological Park, Front Royal, Virginia, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SJC SN AE JG RI BG. Performed the experiments: SN SJC KH JG AE. Analyzed the data: SJC JLB JWC BG. Contributed reagents/materials/analysis tools: KH SBP JLB. Wrote the paper: SJC JLB BG JWC.