About the Authors

Hisatsugu Koshimizu

Affiliations Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Kawaguchi, Japan

Keizo Takao

Affiliations Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Kawaguchi, Japan, Section of Behavior Patterns, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan, Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan

Takashi Matozaki

Affiliations Laboratory of Biosignal Sciences, Institute for Molecular and Cellular Regulation, Gunma University, Maebashi, Japan, Division of Molecular and Cellular Signaling, Department of Biochemistry and Molecular Biology, Kobe University Graduate School of Medicine, Kobe, Japan

Hiroshi Ohnishi

ohnishih@gunma-u.ac.jp (HO); miyakawa@fujita-hu.ac.jp (TM)

Affiliations Laboratory of Biosignal Sciences, Institute for Molecular and Cellular Regulation, Gunma University, Maebashi, Japan, Department of Laboratory Sciences, Gunma University Graduate School of Health Sciences, Maebashi, Japan

Tsuyoshi Miyakawa

ohnishih@gunma-u.ac.jp (HO); miyakawa@fujita-hu.ac.jp (TM)

Affiliations Division of Systems Medical Science, Institute for Comprehensive Medical Science, Fujita Health University, Toyoake, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Kawaguchi, Japan, Section of Behavior Patterns, Center for Genetic Analysis of Behavior, National Institute for Physiological Sciences, Okazaki, Japan, Genetic Engineering and Functional Genomics Group, Frontier Technology Center, Kyoto University Graduate School of Medicine, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HO T. Miyakawa. Performed the experiments: KT HO. Analyzed the data: HK KT HO T. Miyakawa. Contributed reagents/materials/analysis tools: T. Matozaki HO T. Miyakawa. Wrote the paper: HK KT T. Miyakawa.