About the Authors

Hongyu Ma

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China, Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Wei Jiang

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Ping Liu

Affiliation Yellow Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Qingdao, China

Nana Feng

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Qunqun Ma

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Chunyan Ma

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Shujuan Li

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Yuexing Liu

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Zhenguo Qiao

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Lingbo Ma

malingbo@vip.sina.com

Affiliations East China Sea Fisheries Research Institute, Chinese Academy of Fishery Sciences, Shanghai, China, Key Laboratory of East China Sea and Oceanic Fishery Resources Exploitation, Ministry of Agriculture, Shanghai, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HM LM PL. Performed the experiments: HM WJ NF QM SL YL. Analyzed the data: HM WJ. Contributed reagents/materials/analysis tools: HM CM ZQ LM. Wrote the paper: HM. Revised the manuscipt: HM PL.