About the Authors

Mariecia D. Fraser

mariecia.fraser@aber.ac.uk

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom

Jon M. Moorby

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom

James E. Vale

Affiliation Institute of Biological, Environmental and Rural Sciences, Aberystwyth University, Aberystwyth, United Kingdom

Darren M. Evans

Affiliation Department of Biological Sciences, University of Hull, Hull, United Kingdom

Competing Interests

D. Evans has recently served as an Academic Editor for PLOS ONE. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MF JM JV DE. Performed the experiments: MF JM JV DE. Analyzed the data: MF JM JV DE. Wrote the paper: MF JM JV DE.