About the Authors

Liying Feng

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Xue Li

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Qian Yu

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Xianhui Ning

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Jinzhuang Dou

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Jiajun Zou

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Lingling Zhang

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Shi Wang

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Xiaoli Hu

hxl707@ouc.edu.cn (XH); zmbao@ouc.edu.cn (ZB)

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Zhenmin Bao

hxl707@ouc.edu.cn (XH); zmbao@ouc.edu.cn (ZB)

Affiliation Key Laboratory of Marine Genetics and Breeding (MGB), Ministry of Education, College of Marine Life Sciences, Ocean University of China, Qingdao, China

Competing Interests

The authors confirm that the co-author Shi Wang is a PLOS ONE Editorial Board member. This does not alter the authors’ adherence to PLOS ONE Editorial policies and criteria. The authors have also declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ZB XH. Performed the experiments: LF XL QY XN JZ. Analyzed the data: LF JD LZ SW. Contributed reagents/materials/analysis tools: ZB. Wrote the paper: XH LF.