About the Authors

Fengmei Lu

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Yajun Huo

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Meiling Li

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Heng Chen

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Feng Liu

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Yifeng Wang

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Zhiliang Long

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Xujun Duan

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Jiang Zhang

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Ling Zeng

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Huafu Chen

chenhf@uestc.edu.cn

Affiliation Key Laboratory for Neuroinformation of Ministry of Education, School of Life Science and Technology, University of Electronic Science and Technology of China, Chengdu, Sichuan, People’s Republic of China

Competing Interests

The authors state that co-author Huafu Chen is a PLOS ONE Editorial Board member. They declare that no competing interests exist, and this does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: F. Lu YH ML Heng Chen JZ LZ Huafu Chen. Performed the experiments: F. Lu ML Heng Chen F. Liu. Analyzed the data: F. Lu YH Heng Chen YW ZL XD. Contributed reagents/materials/analysis tools: F. Lu YH ML Heng Chen. Wrote the paper: F. Lu YH ML F. Liu.