About the Authors

Matthew J. Pava

Affiliation Department of Neurosciences and Center for Drug and Alcohol Programs, Medical University of South Carolina, Charleston, South Carolina, United States of America

Carolina R. den Hartog

Affiliation Department of Neurosciences and Center for Drug and Alcohol Programs, Medical University of South Carolina, Charleston, South Carolina, United States of America

Carlos Blanco-Centurion

Affiliations Department of Psychiatry & Behavioral Sciences, Medical University of South Carolina, Charleston, South Carolina, United States of America, Ralph H. Johnson VA Medical Center, Charleston, South Carolina, United States of America

Priyattam J. Shiromani

Affiliations Department of Psychiatry & Behavioral Sciences, Medical University of South Carolina, Charleston, South Carolina, United States of America, Ralph H. Johnson VA Medical Center, Charleston, South Carolina, United States of America

John J. Woodward

woodward@musc.edu

Affiliations Department of Neurosciences and Center for Drug and Alcohol Programs, Medical University of South Carolina, Charleston, South Carolina, United States of America, Department of Psychiatry & Behavioral Sciences, Medical University of South Carolina, Charleston, South Carolina, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MJP CRdH CBC PJS JJW. Performed the experiments: MJP CRdH. Analyzed the data: MJP CRdH CBC. Contributed reagents/materials/analysis tools: CBC PJS. Wrote the paper: MJP CRdH JJW.