About the Authors

Zhanghua Chen

zhanghuc@usc.edu (ZC); gross001@umn.edu (MDG); daniel.stram@med.usc.edu (DOS)

Affiliation Department of Preventive Medicine, University of Southern California, Los Angeles, California, United States of America

Mark A. Pereira

Affiliation Division of Epidemiology and Community Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Mark Seielstad

Affiliation Department of Laboratory Medicine, Department of Epidemiology and Biostatistics, and California Institute for Quantitative Biosciences (QB3), University of California San Francisco, San Francisco, California, United States of America

Woon-Puay Koh

Affiliations Duke-National University of Singapore Graduate Medical School, Singapore, Saw Swee Hock School of Public Health, National University of Singapore, Singapore

E. Shyong Tai

Affiliations Duke-National University of Singapore Graduate Medical School, Singapore, Saw Swee Hock School of Public Health, National University of Singapore, Singapore

Yik-Ying Teo

Affiliation Saw Swee Hock School of Public Health, National University of Singapore, Singapore

Jianjun Liu

Affiliations Saw Swee Hock School of Public Health, National University of Singapore, Singapore, Human Genetics, Genome Institute of Singapore, A*STAR, Singapore

Chris Hsu

Affiliation Department of Preventive Medicine, University of Southern California, Los Angeles, California, United States of America

Renwei Wang

Affiliation Department of Epidemiology, University of Pittsburgh Cancer Institute, Pittsburgh, Pennsylvania, United States of America

Andrew O. Odegaard

Affiliation Division of Epidemiology and Community Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Bharat Thyagarajan

Affiliation Division of Epidemiology and Community Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Revati Koratkar

Affiliation Division of Epidemiology and Community Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Jian-Min Yuan

Affiliation Department of Epidemiology, University of Pittsburgh Cancer Institute, Pittsburgh, Pennsylvania, United States of America

Myron D. Gross

zhanghuc@usc.edu (ZC); gross001@umn.edu (MDG); daniel.stram@med.usc.edu (DOS)

Affiliation Division of Epidemiology and Community Health, University of Minnesota, Minneapolis, Minnesota, United States of America

Daniel O. Stram

zhanghuc@usc.edu (ZC); gross001@umn.edu (MDG); daniel.stram@med.usc.edu (DOS)

Affiliation Department of Preventive Medicine, University of Southern California, Los Angeles, California, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: MAP MDG JMY DOS. Performed the experiments: WPK EST YYT JL RW AOO MDG BT RK. Analyzed the data: ZC CH. Wrote the paper: ZC DOS MAP MS MDG.