About the Authors

Hong Liu

Contributed equally to this work with: Hong Liu, Yi Li, Ken Kwok Hon Hung

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Hospital for Skin Diseases, Shandong University, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China, Shandong Provincial Medical Center for Dermatovenereology, Jinan, Shandong, China

Yi Li

Contributed equally to this work with: Hong Liu, Yi Li, Ken Kwok Hon Hung

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Ken Kwok Hon Hung

Contributed equally to this work with: Hong Liu, Yi Li, Ken Kwok Hon Hung

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Na Wang

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Chuan Wang

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Xuechao Chen

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Donglai Sheng

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Xi’an Fu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Kelvin See

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Jia Nee Foo

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Huiqi Low

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Herty Liany

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Ishak Darryl Irwan

Affiliation Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore

Jian Liu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Baoqi Yang

Affiliations Shandong Provincial Hospital for Skin Diseases, Shandong University, Jinan, Shandong, China, Shandong Provincial Medical Center for Dermatovenereology, Jinan, Shandong, China

Mingfei Chen

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Yongxiang Yu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Gongqi Yu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Guiye Niu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Jiabao You

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Yan Zhou

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Shanshan Ma

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Ting Wang

Affiliation Shandong provincial Eye Hospital, Jinan, Shandong, China

Xiaoxiao Yan

Affiliations Shandong Provincial Hospital for Skin Diseases, Shandong University, Jinan, Shandong, China, Shandong Provincial Medical Center for Dermatovenereology, Jinan, Shandong, China

Boon Kee Goh

Affiliation National Skin Centre, Singapore, Singapore

John E. A. Common

Affiliation Institute of Medical Biology, A*STAR, Singapore, Singapore

Birgitte E. Lane

Affiliation Institute of Medical Biology, A*STAR, Singapore, Singapore

Yonghu Sun

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Guizhi Zhou

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Xianmei Lu

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Zhenhua Wang

Affiliation Weifang People’s Hospital, Weifang, Shandong, China

Hongqing Tian

Affiliations Shandong Provincial Hospital for Skin Diseases, Shandong University, Jinan, Shandong, China, Shandong Provincial Medical Center for Dermatovenereology, Jinan, Shandong, China

Yuanhua Cao

Affiliation Institute of Dermatology, Chinese Academy of Medical Sciences and Peking Union Medical College Nanjing, Jiangsu, China

Shumin Chen

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China

Qiji Liu

Affiliation Shandong University, Jinan, Shandong, China

Jianjun Liu

liuj3@gis.a-star.edu.sg (JJL); zhangfuren@hotmail.com (FRZ)

Affiliations Human Genetics, Genome Institute of Singapore, A*STAR, Singapore, Singapore, School of Life Sciences, Anhui Medical University, Hefei, Anhui, China

Furen Zhang

liuj3@gis.a-star.edu.sg (JJL); zhangfuren@hotmail.com (FRZ)

Affiliations Shandong Provincial Institute of Dermatology and Venereology, Provincial Academy of Medical Science, Jinan, Shandong, China, Shandong Provincial Hospital for Skin Diseases, Shandong University, Jinan, Shandong, China, Shandong Provincial Key Lab for Dermatovenereology, Jinan, Shandong, China, Shandong Provincial Medical Center for Dermatovenereology, Jinan, Shandong, China, Shandong Clinical College, Anhui Medical University, Jinan, Shandong, China

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: FZ Jianjun Liu. Performed the experiments: KKHH NW CW XCC DLS XAF KS JNF H. Low H. Liany IDI Jian Liu BQY MFC YXY GQY GYN JBY YZ SSM XXY YHS. Analyzed the data: YL H. Liu. Contributed reagents/materials/analysis tools: TW BKG JEC BEL GZZ XML ZHW HQT YHC SMC QJL. Wrote the paper: H. Liu YL NW.