About the Authors

Jacqueline M. Leung

Affiliations Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America, Program in Cellular, Molecular and Biomedical Sciences, University of Vermont, Burlington, Vermont, United States of America

Mark A. Rould

Affiliation Department of Molecular Physiology and Biophysics, University of Vermont, Burlington, Vermont, United States of America

Christoph Konradt

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Christopher A. Hunter

Affiliation Department of Pathobiology, School of Veterinary Medicine, University of Pennsylvania, Philadelphia, Pennsylvania, United States of America

Gary E. Ward

gary.ward@uvm.edu

Affiliation Department of Microbiology and Molecular Genetics, University of Vermont, Burlington, Vermont, United States of America

Competing Interests

We have the following interests. Gary E. Ward is a member of the Board of Directors of PLOS. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. Mark A. Rould is employed by Macromolecular Consulting LLC. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: JML MAR CK CAH GEW. Performed the experiments: JML CK. Analyzed the data: JML MAR CK CAH GEW. Contributed reagents/materials/analysis tools: JML MAR. Wrote the paper: JML MAR CK CAH GEW.