About the Authors

Chikashi Terao

Contributed equally to this work with: Chikashi Terao, Motomu Hashimoto

a0001101@kuhp.kyoto-u.ac.jp

Affiliation Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Motomu Hashimoto

Contributed equally to this work with: Chikashi Terao, Motomu Hashimoto

Affiliation Department of the Control for Rheumatic Diseases, Kyoto University Graduate School of Medicine, Kyoto, Japan

Moritoshi Furu

Affiliation Department of the Control for Rheumatic Diseases, Kyoto University Graduate School of Medicine, Kyoto, Japan

Shuichiro Nakabo

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Koichiro Ohmura

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Ran Nakashima

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Yoshitaka Imura

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Naoichiro Yukawa

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Hajime Yoshifuji

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Fumihiko Matsuda

Affiliation Center for Genomic Medicine, Kyoto University Graduate School of Medicine, Kyoto, Japan

Hiromu Ito

Affiliation Department of the Control for Rheumatic Diseases, Kyoto University Graduate School of Medicine, Kyoto, Japan

Takao Fujii

Affiliation Department of the Control for Rheumatic Diseases, Kyoto University Graduate School of Medicine, Kyoto, Japan

Tsuneyo Mimori

Affiliation Department of Rheumatology and Clinical Immunology, Kyoto University Graduate School of Medicine, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CT MH FM HI TF TM. Analyzed the data: CT. Contributed reagents/materials/analysis tools: CT MH MF SN KO RN YI NY HY HI TF TM. Wrote the paper: CT MH.