About the Authors

J. Jordan Steel

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Alexander W. E. Franz

Affiliation Department of Veterinary Pathobiology, University of Missouri, Columbia, Missouri, United States of America

Irma Sanchez-Vargas

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Ken E. Olson

Affiliation Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America

Brian J. Geiss

Brian.Geiss@colostate.edu

Affiliations Department of Microbiology, Immunology, and Pathology, Colorado State University, Fort Collins, Colorado, United States of America, Department of Biochemistry and Molecular Biology, Colorado State University, Fort Collins, Colorado, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JJS KEO BJG. Performed the experiments: JJS AWEF IS. Analyzed the data: JJS BJG. Contributed reagents/materials/analysis tools: JJS AWEF IS KEO BJG. Wrote the manuscript: JJS BJG.