About the Authors

Kelly M. McFarlin

kmmcfarlin@alaska.edu

Affiliation Institute of Arctic Biology, University of Alaska Fairbanks, Fairbanks, Alaska, United States of America

Roger C. Prince

Affiliation ExxonMobil Biomedical Sciences, Inc., Annandale, New Jersey, United States of America

Robert Perkins

Affiliation Institute of Northern Engineering, University of Alaska Fairbanks, Fairbanks, Alaska, United States of America

Mary Beth Leigh

Affiliation Institute of Arctic Biology, University of Alaska Fairbanks, Fairbanks, Alaska, United States of America

Competing Interests

As stated in the Financial Disclosure section, this work was part of a Joint Industry Program funded by Shell, ExxonMobil, Statoil, ConocoPhillips and BP. Additional support was provided by the National Institute of General Medical Sciences of the National Institutes of Health under Award Number P20GM103395. This does not alter the authors′ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: KM RCP RP MBL. Performed the experiments: KM RCP RP. Analyzed the data: KM RCP. Contributed reagents/materials/analysis tools: MBL. Wrote the paper: KM RCP RP MBL.