About the Authors

Joanne Preston

Affiliation Biological Sciences, University of Portsmouth, Portsmouth, Hants, United Kingdom

Andrew D. Smith

Current address: Dalton Cumbrian Facility, University of Manchester, Cumbria, United Kingdom

Affiliation STFC Daresbury Laboratory, Warrington, United Kingdom

Eleanor J. Schofield

Affiliation Mary Rose Trust, HM Naval Base, Portsmouth, Hants, United Kingdom

Alan V. Chadwick

Affiliation School of Physical Sciences, University of Kent, Canterbury, Kent, United Kingdom

Mark A. Jones

Affiliation Mary Rose Trust, HM Naval Base, Portsmouth, Hants, United Kingdom

Joy E. M. Watts

joy.watts@port.ac.uk

Affiliation Biological Sciences, University of Portsmouth, Portsmouth, Hants, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JP AS AC ES MJ JW. Performed the experiments: JP AS AC ES. Analyzed the data: JP AS AC ES MJ JW. Contributed reagents/materials/analysis tools: JP AS AC ES MJ JW. Wrote the paper: JP AS AC ES MJ JW.