About the Authors

Anil C. Somenahally

Contributed equally to this work with: Anil C. Somenahally, Jennifer J. Mosher

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Jennifer J. Mosher

Contributed equally to this work with: Anil C. Somenahally, Jennifer J. Mosher

Current address: Stroud Water Research Center, Avondale, Pennsylvania, United States of America

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Tong Yuan

Affiliations Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Institute for Environmental Genomics, Department of Botany, University of Oklahoma, Norman, Oklahoma, United States of America

Mircea Podar

Affiliation ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Tommy J. Phelps

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Joint Biological Sciences Institute, University of Tennessee, Knoxville, Tennessee, United States of America

Steven D. Brown

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Joint Biological Sciences Institute, University of Tennessee, Knoxville, Tennessee, United States of America

Zamin K. Yang

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America

Terry C. Hazen

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Department of Civil & Environmental Engineering, University of Tennessee, Knoxville, Tennessee, United States of America, Department of Earth & Planetary Sciences, University of Tennessee, Knoxville, Tennessee, United States of America, Department of Microbiology, University of Tennessee, Knoxville, Tennessee, United States of America, Joint Biological Sciences Institute, University of Tennessee, Knoxville, Tennessee, United States of America

Adam P. Arkin

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Physical Biosciences Division, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America

Anthony V. Palumbo

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Joint Biological Sciences Institute, University of Tennessee, Knoxville, Tennessee, United States of America

Joy D. Van Nostrand

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Institute for Environmental Genomics, Department of Botany, University of Oklahoma, Norman, Oklahoma, United States of America

Jizhong Zhou

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Institute for Environmental Genomics, Department of Botany, University of Oklahoma, Norman, Oklahoma, United States of America

Dwayne A. Elias

eliasda@ornl.gov

Affiliations ENIGMA – Ecosystems and Networks Integrated with Genes and Molecular Assemblies Program, Lawrence Berkeley National Laboratory, Berkeley, California, United States of America, Biosciences Division, Oak Ridge National Laboratory, Oak Ridge, Tennessee, United States of America, Joint Biological Sciences Institute, University of Tennessee, Knoxville, Tennessee, United States of America

Competing Interests

Dr. Elias is a PLOS ONE Editorial Board member. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: ACS JJM MP TJP SDB AVP DAE. Performed the experiments: ACS JJM ZKY. Analyzed the data: ACS JJM TY JDVN JZ DAE. Contributed reagents/materials/analysis tools: DAE TCH APA. Wrote the paper: ACS JJM DAE.