About the Authors

Eugene Y. Chiang

Affiliation Department of Immunology, Genentech Inc., South San Francisco, California, United States of America

Robert J. Johnston

Affiliation Department of Immunology, Genentech Inc., South San Francisco, California, United States of America

Jane L. Grogan

jgrogan@gene.com

Affiliation Department of Immunology, Genentech Inc., South San Francisco, California, United States of America

Competing Interests

All authors are employees of Genentech, a member of the Roche group, which develops and markets drugs for profit. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: EC JG. Performed the experiments: EC RJ. Analyzed the data: EC RJ JG. Wrote the paper: EC RJ JG.