About the Authors

James D. Foster

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush, Midlothian, Scotland, United Kingdom

Wilfred Goldmann

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush, Midlothian, Scotland, United Kingdom

Nora Hunter

nora.hunter@roslin.ed.ac.uk

Affiliation The Roslin Institute and Royal (Dick) School of Veterinary Studies, The University of Edinburgh, Easter Bush, Midlothian, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: JDF WG NH. Performed the experiments: JDF WG NH. Analyzed the data: JDF NH. Contributed reagents/materials/analysis tools: JDF WG NH. Wrote the paper: JDF WG NH.