About the Authors

Hiromi Hyodo

Affiliation University of Tsukuba, Faculty of Life and Environmental Sciences, Tsukuba, Ibaraki, Japan

Azusa Terao

Affiliation University of Tsukuba, Faculty of Life and Environmental Sciences, Tsukuba, Ibaraki, Japan

Jun Furukawa

Affiliation University of Tsukuba, Faculty of Life and Environmental Sciences, Tsukuba, Ibaraki, Japan

Naoya Sakamoto

Affiliation Hokkaido University, Creative Research Institution (CRIS), Sapporo, Hokkaido, Japan

Hisayoshi Yurimoto

Affiliations Hokkaido University, Creative Research Institution (CRIS), Sapporo, Hokkaido, Japan, Hokkaido University, Natural History Sciences, Sapporo, Hokkaido, Japan

Shinobu Satoh

Affiliation University of Tsukuba, Faculty of Life and Environmental Sciences, Tsukuba, Ibaraki, Japan

Hiroaki Iwai

iwai.hiroaki.gb@u.tsukuba.ac.jp

Affiliation University of Tsukuba, Faculty of Life and Environmental Sciences, Tsukuba, Ibaraki, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HI. Performed the experiments: HH. Analyzed the data: HH AT JF SS HI. Contributed reagents/materials/analysis tools: NS HY. Wrote the paper: HH HI.