About the Authors

Rebecca L. Fagan

Affiliation Department of Biochemistry, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Meng Wu

Affiliation Department of Biochemistry, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Frédéric Chédin

Affiliation Department of Molecular and Cellular Biology and Genome Center, University of California Davis, Davis, California, United States of America

Charles Brenner

charles-brenner@uiowa.edu

Affiliation Department of Biochemistry, Carver College of Medicine, University of Iowa, Iowa City, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RLF MW CB. Performed the experiments: RLF. Analyzed the data: RLF MW CB. Contributed reagents/materials/analysis tools: MW FC. Wrote the paper: RLF CB.