About the Authors

Nicolai A. Kittan

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Ronald M. Allen

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Abhay Dhaliwal

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Karen A. Cavassani

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Matthew Schaller

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Katherine A. Gallagher

Affiliation Division of Vascular Surgery, University of Michigan Hospital, Ann Arbor, Michigan, United States of America

William F. Carson IV

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Sumanta Mukherjee

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Jolanta Grembecka

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Tomasz Cierpicki

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Gabor Jarai

Affiliation Novartis Institutes of Biomedical Research, Respiratory Disease Area, Horsham, West Sussex, United Kingdom

John Westwick

Affiliation Novartis Institutes of Biomedical Research, Respiratory Disease Area, Horsham, West Sussex, United Kingdom

Steven L. Kunkel

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Cory M. Hogaboam

hogaboam@med.umich.edu

Affiliation Department of Pathology, University of Michigan Medical School, Ann Arbor, Michigan, United States of America

Competing Interests

The authors confirm that they will fully adhere to all the PLOS ONE policies. They received funding from Novartis. And two of the coauthors, Gabor Jarai and John Westwick, are (GJ) or were (JW) employed by the Novartis Institutes for Biomedical Research, UK. However, this does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials. The authors also have no other commercial interests to declare.

Author Contributions

Conceived and designed the experiments: NAK RMA KAC MS KAG WFC SM JG TC GJ JW SLK CMH. Performed the experiments: NAK RMA AD KAC MS SM. Analyzed the data: NAK RMA KAC MS KAG WFC SM JG TC SLK CMH. Contributed reagents/materials/analysis tools: JG TC. Wrote the manuscript: NAK RMA SLK CMH.