About the Authors

Zhi-Hua Li

Affiliations Department of Biological Sciences, National University of Singapore, Singapore, Yangtze River Fisheries Research Institute, Chinese Academy of Fishery Sciences, Wuhan, China

Hongyan Xu

Affiliation Department of Biological Sciences, National University of Singapore, Singapore

Weiling Zheng

Affiliation Department of Biological Sciences, National University of Singapore, Singapore

Siew Hong Lam

Affiliation Department of Biological Sciences, National University of Singapore, Singapore

Zhiyuan Gong

dbsgzy@nus.edu.sg

Affiliation Department of Biological Sciences, National University of Singapore, Singapore

Competing Interests

Zhiyuan Gong is an Academic Editor of PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: HX SHL ZG. Performed the experiments: HX. Analyzed the data: ZHL WZ ZG. Contributed reagents/materials/analysis tools: ZHL HX WZ SHL. Wrote the paper: ZHL ZG.