About the Authors

Nadia Zaraî Jaouadi

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Hatem Rekik

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Abdelmalek Badis

Affiliations Laboratory of Natural Products Chemistry and Biomolecules (LNPCB), University of Saàd Dahlab (USD) of Blida, Blida, Algeria, National Centre for Research and Development of Fisheries and Aquaculture (CNRDPA), Bousmail (W. Tipaza), Algeria

Sahar Trabelsi

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Mouna Belhoul

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Amina Benkiar Yahiaoui

Affiliation National Centre for Research and Development of Fisheries and Aquaculture (CNRDPA), Bousmail (W. Tipaza), Algeria

Houda Ben Aicha

Affiliation National Leather and Shoe Center (CNCC), Mégrine, Ben Arous, Tunisia

Abdessatar Toumi

Affiliation National Leather and Shoe Center (CNCC), Mégrine, Ben Arous, Tunisia

Samir Bejar

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Bassem Jaouadi

bassem.jaouadi@yahoo.fr

Affiliation Laboratory of Microorganisms and Biomolecules (LMB), Centre of Biotechnology of Sfax (CBS), University of Sfax, Sfax, Tunisia

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: BJ SB. Performed the experiments: NZJ HR ST MB. Analyzed the data: NZJ AB ABY HBA AT. Contributed reagents/materials/analysis tools: HBA AT. Wrote the paper: NZJ BJ SB. Critical revision of manuscript: NZJ BJ SB.