About the Authors

Yali Hou

Contributed equally to this work with: Yali Hou, Yachun Wang, Xuemei Lu

Affiliations Key Laboratory of Agricultural Animal Genetics and Breeding, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, People’s Republic of China, Laboratory of Disease Genomics and Individualized Medicine, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, People’s Republic of China

Yachun Wang

Contributed equally to this work with: Yali Hou, Yachun Wang, Xuemei Lu

Affiliation Key Laboratory of Agricultural Animal Genetics and Breeding, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, People’s Republic of China

Xuemei Lu

Contributed equally to this work with: Yali Hou, Yachun Wang, Xuemei Lu

Affiliation Laboratory of Disease Genomics and Individualized Medicine, Beijing Institute of Genomics, Chinese Academy of Sciences, Beijing, People’s Republic of China

Xu Zhang

Affiliation Key Laboratory of Agricultural Animal Genetics and Breeding, National Engineering Laboratory for Animal Breeding, College of Animal Science and Technology, China Agricultural University, Beijing, People’s Republic of China

Qian Zhao

Affiliation College of Animal Science and Technology, Northeast Agricultural University, Harbin, People’s Republic of China

Rory J. Todhunter

rjt2@cornell.edu (RJT); zz19@cornell.edu (ZZ)

Affiliation Department of Clinical Sciences, College of Veterinary Medicine, Cornell University, Ithaca, New York, United States of America

Zhiwu Zhang

rjt2@cornell.edu (RJT); zz19@cornell.edu (ZZ)

Affiliations College of Animal Science and Technology, Northeast Agricultural University, Harbin, People’s Republic of China, Institute for Genomic Diversity, Cornell University, Ithaca, New York, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: RJT ZZ. Performed the experiments: YH. Analyzed the data: YH. Contributed reagents/materials/analysis tools: YW XL XZ QZ. Wrote the paper: YH YW XL RJT ZZ.