About the Authors

Xu Xu

Affiliation Department of Biology, Center for Cellular Dynamics and the Huck Institute of the Life Sciences, Penn State University, University Park, Pennsylvania, United States of America

Jingjie Hu

Affiliation Department of Biology, Center for Cellular Dynamics and the Huck Institute of the Life Sciences, Penn State University, University Park, Pennsylvania, United States of America

Barbara C. McGrath

Affiliation Department of Biology, Center for Cellular Dynamics and the Huck Institute of the Life Sciences, Penn State University, University Park, Pennsylvania, United States of America

Douglas R. Cavener

drc9@psu.edu

Affiliation Department of Biology, Center for Cellular Dynamics and the Huck Institute of the Life Sciences, Penn State University, University Park, Pennsylvania, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: XX JH BCM DRC. Performed the experiments: XX JH DRC. Analyzed the data: XX JH BCM DRC. Wrote the paper: XX JH BCM DRC.