About the Authors

Tatsuro Kamijima

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan

Mami Sakashita

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan

Ai Miura

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan

Yoshiyuki Nishimiya

Affiliation Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan

Sakae Tsuda

s.tsuda@aist.go.jp

Affiliations Bioproduction Research Institute, National Institute of Advanced Industrial Science and Technology (AIST), Sapporo, Hokkaido, Japan, Graduate School of Life Science, Hokkaido University, Sapporo, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK ST. Performed the experiments: TK MS AM YN. Analyzed the data: TK YN. Contributed reagents/materials/analysis tools: TK YN. Wrote the paper: TK ST.