About the Authors

Takayuki Fujiwara

Current address: National Institute of Genetics, Mishima, Japan

Affiliation Chromosome Dynamics Laboratory, RIKEN, Wako, Saitama, Japan

Mio Ohnuma

Affiliations Faculty of Science, Rikkyo University, Toshima-ku, Tokyo, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Chiyoda-ku, Tokyo, Japan

Masaki Yoshida

Affiliation Faculty of Life & Environmental Sciences, University of Tsukuba, Tsukuba, Ibaraki, Japan

Tsuneyoshi Kuroiwa

Affiliations Faculty of Science, Rikkyo University, Toshima-ku, Tokyo, Japan, Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Chiyoda-ku, Tokyo, Japan

Tatsuya Hirano

hiranot@riken.jp

Affiliation Chromosome Dynamics Laboratory, RIKEN, Wako, Saitama, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TF TH. Performed the experiments: TF. Analyzed the data: TF TH. Contributed reagents/materials/analysis tools: MO MY TK. Wrote the paper: TF TH.