About the Authors

Tracy L. Nicholson

tracy.nicholson@ars.usda.gov

Affiliation National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Sarah M. Shore

Affiliation National Animal Disease Center, Agricultural Research Service, United States Department of Agriculture, Ames, Iowa, United States of America

Tara C. Smith

Affiliation Center for Emerging Infectious Diseases, Department of Epidemiology, College of Public Health, University of Iowa, Iowa City, Iowa, United States of America

Timothy S. Fraena

Affiliation Department of Veterinary Diagnostic and Production Animal Medicine, College of Veterinary Medicine, Iowa State University, Ames, Iowa, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TLN. Performed the experiments: SMS. Analyzed the data: TLN SMS. Contributed reagents/materials/analysis tools: TCS TSF. Wrote the manuscript: TLN SMS. Critically reviewed manuscript: TLN SMS TCS TSF.