About the Authors

Kumi Kaneko

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Tsubomi Ikeda

Contributed equally to this work with: Tsubomi Ikeda, Mirai Nagai, Sayaka Hori

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Mirai Nagai

Contributed equally to this work with: Tsubomi Ikeda, Mirai Nagai, Sayaka Hori

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Sayaka Hori

Contributed equally to this work with: Tsubomi Ikeda, Mirai Nagai, Sayaka Hori

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Chie Umatani

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Hiroto Tadano

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Atsushi Ugajin

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Takayoshi Nakaoka

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Rajib Kumar Paul

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Tomoko Fujiyuki

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Kenichi Shirai

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Takekazu Kunieda

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Hideaki Takeuchi

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Takeo Kubo

stkubo@biol.s.u-tokyo.ac.jp

Affiliation Department of Biological Sciences, Graduate School of Science, The University of Tokyo, Bunkyo-ku, Tokyo, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KK TI MN SH CU H. Tadano AU T. Kunieda H. Takeuchi. Performed the experiments: KK TI MN SH CU. Analyzed the data: KK TI MN SH CU H. Tadano AU. Contributed reagents/materials/analysis tools: TN RKP TF KS. Wrote the paper: KK T. Kubo.