About the Authors

Saija J. Ahonen

Affiliations Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland

Elina Pietilä

Affiliation Department of Equine and Small Animal Medicine, Faculty of Veterinary Medicine, University of Helsinki, Helsinki, Finland

Cathryn S. Mellersh

Affiliation Canine Genetics Animal Health Trust, Lanwades Park, Kentford, Newmarket, Suffolk, United Kingdom

Katriina Tiira

Affiliations Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland

Liz Hansen

Affiliation Department of Veterinary Pathobiology, College of Veterinary Medicine, University of Missouri, Missouri, United States of America

Gary S. Johnson

Affiliation Department of Veterinary Pathobiology, College of Veterinary Medicine, University of Missouri, Missouri, United States of America

Hannes Lohi

hannes.lohi@helsinki.fi

Affiliations Department of Veterinary Biosciences and Research Programs Unit, Molecular Neurology, University of Helsinki, Helsinki, Finland, The Folkhälsan Institute of Genetics, Helsinki, Finland

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: SA EP KT HL. Performed the experiments: SA EP. Analyzed the data: SA EP KT. Contributed reagents/materials/analysis tools: CM GJ LH HL. Wrote the paper: SA EP HL.