About the Authors

Natsuko Tsujino

Affiliations Department of Molecular Neuroscience and Integrative Physiology, Faculty of Medicine, Kanazawa University, Kanazawa, Ishikawa, Japan, International Institute for Integrative Sleep Medicine, University of Tsukuba, Tsukuba, Japan

Tomomi Tsunematsu

Affiliation Research Institute of Environmental Medicine, Nagoya University, Nagoya, Japan

Motokazu Uchigashima

Affiliation Department of Anatomy and Embryology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Kohtarou Konno

Affiliation Department of Anatomy and Embryology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Akihiro Yamanaka

Affiliation Research Institute of Environmental Medicine, Nagoya University, Nagoya, Japan

Kazuto Kobayashi

Affiliation Department of Molecular Genetics, Fukushima Medical University School of Medicine, Fukushima, Japan

Masahiko Watanabe

Affiliation Department of Anatomy and Embryology, Graduate School of Medicine, Hokkaido University, Sapporo, Japan

Yoshimasa Koyama

tsakurai@med.kanazawa-u.ac.jp (TS); koyamay@sss.fukushima-u.ac.jp (YK)

Affiliation Department of Science and Technology, Fukushima University, Fukushima, Japan

Takeshi Sakurai

tsakurai@med.kanazawa-u.ac.jp (TS); koyamay@sss.fukushima-u.ac.jp (YK)

Affiliations Department of Molecular Neuroscience and Integrative Physiology, Faculty of Medicine, Kanazawa University, Kanazawa, Ishikawa, Japan, International Institute for Integrative Sleep Medicine, University of Tsukuba, Tsukuba, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YK TS. Performed the experiments: NT TT MU K. Konno. Analyzed the data: NT AY MW TS. Contributed reagents/materials/analysis tools: K. Kobayashi. Wrote the paper: NT TS.