About the Authors

Robert Cook

Affiliations School of Ocean Sciences, Bangor University, Menai Bridge, Anglesey, United Kingdom, School of Life Sciences, Heriot-Watt University, Edinburgh, United Kingdom

Jose M. Fariñas-Franco

Affiliation School of Life Sciences, Heriot-Watt University, Edinburgh, United Kingdom

Fiona R. Gell

Affiliation Fisheries Directorate, Department of Environment Food and Agriculture, Isle of Man Government, St John's, Isle of Man, United Kingdom

Rohan H. F. Holt

Affiliation Natural Resources Wales, Bangor, Gwynedd, United Kingdom

Terry Holt

Affiliation Center for Marine and Coastal Studies Ltd. (CMACS), Port Erin, Isle of Man, United Kingdom

Charles Lindenbaum

Affiliation Natural Resources Wales, Bangor, Gwynedd, United Kingdom

Joanne S. Porter

Affiliation School of Life Sciences, Heriot-Watt University, Edinburgh, United Kingdom

Ray Seed

Affiliation School of Ocean Sciences, Bangor University, Menai Bridge, Anglesey, United Kingdom

Lucie R. Skates

Affiliation Natural Resources Wales, Bangor, Gwynedd, United Kingdom

Thomas B. Stringell

Affiliation Natural Resources Wales, Bangor, Gwynedd, United Kingdom

William G. Sanderson

w.g.sanderson@hw.ac.uk

Affiliations Natural Resources Wales, Bangor, Gwynedd, United Kingdom, School of Life Sciences, Heriot-Watt University, Edinburgh, United Kingdom

Competing Interests

The authors have the following interests: Terry Holt is employed by CMACS Ltd. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: RLC CL FRG RHFH JSP TH RS LRS TBS WGS. Performed the experiments: RLC RHFH CL JSP TBS WGS. Analyzed the data: RLC JMF LRS TBS WGS. Wrote the paper: RLC CL FRG WGS JMF TH RS TBS JSP. Logistics in the Isle of Man: FRG TH .