About the Authors

Yusuke Murata

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Kyoji Nishio

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Takayuki Mochiyama

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Morichika Konishi

mkonishi@kobepharma-u.ac.jp (MK); itohnobu@pharm.kyoto-u.ac.jp (NI)

Affiliations Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan, Department of Microbial Chemistry, Kobe Pharmaceutical University, Motoyamakita-machi, Kobe, Hyogo, Japan

Masaya Shimada

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Hiroya Ohta

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Nobuyuki Itoh

mkonishi@kobepharma-u.ac.jp (MK); itohnobu@pharm.kyoto-u.ac.jp (NI)

Affiliation Department of Genetic Biochemistry, Kyoto University Graduate School of Pharmaceutical Sciences, Sakyo, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: YM MK HO NI. Performed the experiments: YM KN TM MK MS. Analyzed the data: YM KN MK MS HO NI. Wrote the paper: YM MK NI.