About the Authors

Brent J. Sewall

bjsewall@temple.edu

Affiliations Department of Biology, Temple University, Philadelphia, Pennsylvania, United States of America, Department of Wildlife, Fish, and Conservation Biology, University of California Davis, Davis, California, United States of America

Amy L. Freestone

Affiliation Department of Biology, Temple University, Philadelphia, Pennsylvania, United States of America

Joseph E. Hawes

Affiliation School of Environmental Sciences, University of East Anglia, Norwich Research Park, Norwich, United Kingdom

Ernest Andriamanarina

Affiliation Département des Sciences de la Nature et de l'Environnement, Université d'Antsiranana, Antsiranana, Madagascar

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Analyzed the data: BJS. Wrote the paper: BJS. Conceived of project: BJS. Developed conceptual framework: BJS ALF. Designed observations: BJS ALF JEH EA. Managed field teams: BJS JEH. Collected data: BJS ALF JEH EA. Edited the manuscript: BJS ALF JEH.