About the Authors

Peter K. Joshi

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

James Prendergast

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Ross M. Fraser

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Jennifer E. Huffman

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Veronique Vitart

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Caroline Hayward

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Ruth McQuillan

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Dominik Glodzik

Affiliations Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom, MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Ozren Polašek

Affiliations Department of Public Health, University of Split, Split, Croatia, Centre for Global Health, University of Split, Split, Croatia

Nicholas D. Hastie

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Igor Rudan

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Harry Campbell

Affiliation Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Alan F. Wright

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Chris S. Haley

Affiliations MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom, Roslin Institute, University of Edinburgh, Scotland, United Kingdom

James F. Wilson

Contributed equally to this work with: James F. Wilson, Pau Navarro

pau.navarro@ed.ac.uk (PN); jim.wilson@ed.ac.uk (JFW)

Affiliations Centre for Population Health Sciences, University of Edinburgh, Edinburgh, Scotland, United Kingdom, MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Pau Navarro

Contributed equally to this work with: James F. Wilson, Pau Navarro

pau.navarro@ed.ac.uk (PN); jim.wilson@ed.ac.uk (JFW)

Affiliation MRC Human Genetics Unit, University of Edinburgh, Edinburgh, Scotland, United Kingdom

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: CSH JFW PN. Performed the experiments: PKJ. Analyzed the data: PKJ JP RFM. Contributed reagents/materials/analysis tools: JEH VV CH RM DG OP NDH IR HC AFW CSH JFW PN. Wrote the paper: PKJ CSH JFW PN.