About the Authors

Claude Loverdo

loverdo@ucla.edu

Affiliation Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, California, United States of America

James O. Lloyd-Smith

Affiliations Department of Ecology and Evolutionary Biology, University of California Los Angeles, Los Angeles, California, United States of America, Fogarty International Center, National Institutes of Health, Bethesda, Maryland, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Wrote the paper: CL JLS. Conceived and designed the theory and calculations: CL JLS. Performed the calculations: CL. Analyzed the results from theory and calculations: CL.