About the Authors

Koh Nomura

Affiliation Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan

Takahiro Yonezawa

cyclotis@gmail.com

Affiliations School of Life Sciences, Fudan University, Shanghai, China, Institute of the Statistical Mathematics, Tokyo, Japan

Shuhei Mano

Affiliation Institute of the Statistical Mathematics, Tokyo, Japan

Shigehisa Kawakami

Affiliation Gunma Safari World co. Ltd, Gunma, Japan

Andrew M. Shedlock

Affiliation College of Charleston Department of Biology and Medical University of South Carolina College of Graduate Studies, Charleston, South Carolina, United States of America

Masami Hasegawa

Affiliations School of Life Sciences, Fudan University, Shanghai, China, Institute of the Statistical Mathematics, Tokyo, Japan

Takashi Amano

Affiliation Faculty of Agriculture, Tokyo University of Agriculture, Kanagawa, Japan

Competing Interests

Shigehisa Kawakami is employed by Gunma Safari World co. Ltd. Blood samples of bezoar were collected from Gunma Safari World Co. Ltd, Japan for this study. There are no patents, products in development or marketed products to declare. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: KN TY MH TA. Performed the experiments: KN TA. Analyzed the data: TY SM MH. Contributed reagents/materials/analysis tools: SK. Wrote the paper: KN TY SM AMS MH.