About the Authors

Anna R. Holden

aholden@nhm.org

Affiliation Department of Entomology, Natural History Museum of Los Angeles County, Los Angeles, California, United States of America

John M. Harris

Affiliation George C. Page Museum, Los Angeles, California, United States of America

Robert M. Timm

Affiliation Department of Ecology and Evolutionary Biology and Biodiversity Institute, University of Kansas, Lawrence, Kansas, United States of America

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: ARH RMT. Performed the experiments: ARH RMT. Analyzed the data: ARH. Contributed reagents/materials/analysis tools: JMH. Wrote the paper: ARH JMH RMT.