About the Authors

Masaki Shoji

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Etsuhisa Takahashi

Affiliation Division of Enzyme Chemistry, Institute for Enzyme Research, The University of Tokushima, Tokushima, Japan

Dai Hatakeyama

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Yuma Iwai

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Yuka Morita

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Riku Shirayama

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Noriko Echigo

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Hiroshi Kido

Affiliation Division of Enzyme Chemistry, Institute for Enzyme Research, The University of Tokushima, Tokushima, Japan

Shigeo Nakamura

Affiliation Department of Chemistry, Nippon Medical School, Nakahara-ku, Kawasaki, Kanagawa, Japan

Tadahiko Mashino

Affiliation Department of Pharmaceutical Sciences, Faculty of Pharmacy, Keio University, Minato-ku, Tokyo, Japan

Takeshi Okutani

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Takashi Kuzuhara

kuzuhara@ph.bunri-u.ac.jp

Affiliation Laboratory of Biochemistry, Faculty of Pharmaceutical Sciences, Tokushima Bunri University, Yamashiro-cho, Tokushima, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: TK. Performed the experiments: MS ET DH YI YM RS NE TO. Analyzed the data: MS ET YI YM RS HK. Contributed reagents/materials/analysis tools: SN TM ET HK. Wrote the paper: TK MS.