About the Authors

Bae Hyun Shin

Affiliation Global Research Laboratory, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Yunki Lim

Affiliation College of Life Sciences, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Hye Jin Oh

Affiliation College of Life Sciences, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Sang Min Park

Affiliation Global Research Laboratory, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Sun-Kyung Lee

Affiliations Department of Life Science, Hanyang University, Seoul, Korea, The Research Institute for Natural Sciences, Hanyang University, Seoul, Korea

Joohong Ahnn

Affiliation Department of Life Science, Hanyang University, Seoul, Korea

Do Han Kim

Affiliation College of Life Sciences, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Woo Keun Song

Affiliation College of Life Sciences, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Tae Hwan Kwak

Affiliation R&D Institute, Mazence Inc., Suwon, Korea

Woo Jin Park

woojinpark@me.com

Affiliation Global Research Laboratory, Gwangju Institute of Science and Technology (GIST), Gwangju, Korea

Competing Interests

Tae Hwan Kwak is an employee of R&D Mazence Inc. and has a patent which protects the use of β-lapachone for the treatment of metabolic diseases. This does not alter the authors’ adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: BHS WJP. Performed the experiments: BHS YL HJO SMP. Analyzed the data: BHS YL HJO SKL. Contributed reagents/materials/analysis tools: JA DHK WKS THK. Wrote the paper: BHS WJP.