About the Authors

Rajapandian SivaGanesa Karthikeyan

Affiliation Dr. G. Venkatasamy Eye Research Institute, Aravind Eye Hospital, Madurai, Tamil Nadu, India

Jeganathan Lakshmi Priya

Affiliation Dr. G. Venkatasamy Eye Research Institute, Aravind Eye Hospital, Madurai, Tamil Nadu, India

Sixto M. Leal Jr

Affiliation Department of Ophthalmology and Visual Sciences, Case Western Reserve University, Cleveland, Ohio, United States of America

Jonida Toska

Affiliation Department of Molecular Biology and Microbiology, Case Western Reserve University, Cleveland, Ohio, United States of America

Arne Rietsch

Affiliation Department of Molecular Biology and Microbiology, Case Western Reserve University, Cleveland, Ohio, United States of America

Venkatesh Prajna

Affiliation Dr. G. Venkatasamy Eye Research Institute, Aravind Eye Hospital, Madurai, Tamil Nadu, India

Eric Pearlman

Affiliation Department of Ophthalmology and Visual Sciences, Case Western Reserve University, Cleveland, Ohio, United States of America

Prajna Lalitha

lalitha@aravind.org

Affiliation Dr. G. Venkatasamy Eye Research Institute, Aravind Eye Hospital, Madurai, Tamil Nadu, India

Competing Interests

We have the following interest. EP is a recipient of an Alcon Research Institute award. There are no patents, products in development or marketed products to declare. This does not alter our adherence to all the PLOS ONE policies on sharing data and materials, as detailed online in the guide for authors.

Author Contributions

Conceived and designed the experiments: RSK SML EP AR VP PL. Performed the experiments: RSK JLP SML JT. Analyzed the data: RSK SML EP AR VP PL. Wrote the paper: RSK SML EP AR VP PL.