About the Authors

Jun-ichi Kishikawa

Affiliation Department of Molecular Biosciences, Kyoto Sangyo University, Motoyama Kamigamo, Kita-ku, Kyoto, Japan

Tatsuya Ibuki

Affiliation Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Shuichi Nakamura

Affiliations Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan, Department of Applied Physics, Graduate School of Engineering, Tohoku University, Sendai, Miyagi, Japan

Astuko Nakanishi

Affiliation Department of Molecular Biosciences, Kyoto Sangyo University, Motoyama Kamigamo, Kita-ku, Kyoto, Japan

Tohru Minamino

Affiliations Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan, Precursory Research for Embryonic Science and Technology, Japan Science and Technology Agency, Kawaguchi, Saitama, Japan

Tomoko Miyata

Affiliation Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan

Keiichi Namba

Affiliations Graduate School of Frontier Biosciences, Osaka University, Osaka, Japan, Riken Quantitative Biology Center, Osaka, Japan

Hiroki Konno

Affiliation Imaging Research Division, Bio-AFM Frontier Research Center, Kanazawa University, Kanazawa, Ishikawa, Japan

Hiroshi Ueno

Affiliation Department of Physics, Faculty of Science and Engineering, Chuo University, Bunkyo-ku, Tokyo, Japan

Katsumi Imada

kimada@chem.sci.osaka-u.ac.jp (KI); yokoken@cc.kyoto-su.ac.jp (KY)

Affiliation Graduate School of Science, Osaka University, Toyonaka, Osaka, Japan

Ken Yokoyama

kimada@chem.sci.osaka-u.ac.jp (KI); yokoken@cc.kyoto-su.ac.jp (KY)

Affiliation Department of Molecular Biosciences, Kyoto Sangyo University, Motoyama Kamigamo, Kita-ku, Kyoto, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KY KI. Performed the experiments: JK IT AN SN HU HK. Analyzed the data: JK IT AN SN HU HK. Contributed reagents/materials/analysis tools: KN. Wrote the paper: KY KI TM.