About the Authors

Qingguo Wang

Affiliation Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Peilin Jia

Affiliations Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Center for Quantitative Sciences, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Zhongming Zhao

zhongming.zhao@vanderbilt.edu

Affiliations Department of Biomedical Informatics, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Psychiatry, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Department of Cancer Biology, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America, Center for Quantitative Sciences, Vanderbilt University School of Medicine, Nashville, Tennessee, United States of America

Competing Interests

The authors have the following interest. Zhongming Zhao has served as an editorial board member of PLOS ONE since 2010. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: QW ZZ. Performed the experiments: QW. Analyzed the data: QW PJ ZZ. Contributed reagents/materials/analysis tools: QW. Wrote the paper: QW PJ ZZ.