About the Authors

Megan Y. Murray

Affiliation School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk, United Kingdom

Timothy P. Birkland

Affiliation Center for Lung Biology, University of Washington, Seattle, Washington, United States of America

Jonathan D. Howe

Current address: MRC Laboratory of Molecular Biology, Cambridge United Kingdom

Affiliation School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk, United Kingdom

Andrew D. Rowan

Affiliation Musculoskeletal Research Group, Institute of Cellular Medicine, The Medical School, Newcastle University, Newcastle, United Kingdom

Mark Fidock

Affiliation Pfizer Global Research and Development, Sandwich, Kent, United Kingdom

William C. Parks

Affiliation Center for Lung Biology, University of Washington, Seattle, Washington, United States of America

Jelena Gavrilovic

J.Gavrilovic@uea.ac.uk

Affiliation School of Biological Sciences, University of East Anglia, Norwich Research Park, Norwich, Norfolk, United Kingdom

Competing Interests

Dr Mark Fidock was an employee of Pfizer Global Research and Development. The authors confirm that this does not alter their adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: MYM MF WCP JG. Performed the experiments: MYM TPB JDH. Analyzed the data: MYM JDH JG. Contributed reagents/materials/analysis tools: ADR WCP. Wrote the paper: MYM WCP JG. Directed the research: JG.