About the Authors

Stephen C. McDowell

Affiliation Department of Biochemistry and Molecular Biology, University of Nevada, Reno, Nevada, United States of America

Garo Akmakjian

Affiliation Division of Biological Sciences, University of California, San Diego, California, United States of America

Chris Sladek

Affiliation Department of Biochemistry and Molecular Biology, University of Nevada, Reno, Nevada, United States of America

David Mendoza-Cozatl

Affiliation Division of Biological Sciences, University of California, San Diego, California, United States of America

Joe B. Morrissey

Affiliation Department of Biological Sciences, Dartmouth College, Hanover, New Hampshire, United States of America

Nick Saini

Affiliation Department of Biochemistry and Molecular Biology, University of Nevada, Reno, Nevada, United States of America

Ron Mittler

Affiliation Department of Biological Sciences, University of North Texas, Denton, Texas, United States of America

Ivan Baxter

Affiliation United States Department of Agriculture–Agricultural Research Service, Plant Genetics Research Unit, Donald Danforth Plant Science Center, St. Louis, Missouri, United States of America

David E. Salt

Affiliation Institute of Biological and Environmental Science, University of Aberdeen, Aberdeen, Scotland, United Kingdom

John M. Ward

Affiliation Department of Plant Biology, University of Minnesota, St. Paul, Minnesota, United States of America

Julian I. Schroeder

Affiliation Division of Biological Sciences, University of California, San Diego, California, United States of America

Mary Lou Guerinot

Affiliation Department of Biological Sciences, Dartmouth College, Hanover, New Hampshire, United States of America

Jeffrey F. Harper

jfharper@unr.edu.

Affiliation Department of Biochemistry and Molecular Biology, University of Nevada, Reno, Nevada, United States of America

Competing Interests

It should be noted that Ivan Baxter is a co-author of this manuscript and is also an editor for PLOS ONE. This does not alter the authors' adherence to all the PLOS ONE policies on sharing data and materials.

Author Contributions

Conceived and designed the experiments: SCM RM IB DS JW JIS MLG JFH. Performed the experiments: SCM GA CS DMC JBM NS JFH JW. Analyzed the data: SCM GA CS IB DS JW JIS MLG JFH. Contributed reagents/materials/analysis tools: SCM JW RM IB DS JIS MLG JFH. Wrote the paper: SCM GA JFH IB DS MLG JIS JBM JW.