About the Authors

Tzu-Yen Chen

Contributed equally to this work with: Tzu-Yen Chen, Da-Yuan Chen

Affiliation Department of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Da-Yuan Chen

Contributed equally to this work with: Tzu-Yen Chen, Da-Yuan Chen

Current address: Department of Microbiology & Immunology, University of Otago, Otago, New Zealand

Affiliation Graduate Institute of Microbiology and Public Health, National Chung Hsing University, Taichung, Taiwan

Hsiao-Wei Wen

Affiliation Department of Food Science and Biotechnology, National Chung Hsing University, Taichung, Taiwan

Jun-Lin Ou

Affiliation Graduate Institute of Microbiology and Public Health, National Chung Hsing University, Taichung, Taiwan

Shyan-Song Chiou

Affiliation Graduate Institute of Microbiology and Public Health, National Chung Hsing University, Taichung, Taiwan

Jo-Mei Chen

Affiliation Graduate Institute of Microbiology and Public Health, National Chung Hsing University, Taichung, Taiwan

Min-Liang Wong

Affiliation Department of Veterinary Medicine, National Chung Hsing University, Taichung, Taiwan

Wei-Li Hsu

wlhsu@dragon.nchu.edu.tw

Affiliation Graduate Institute of Microbiology and Public Health, National Chung Hsing University, Taichung, Taiwan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: HWW SSC MLW WLH. Performed the experiments: TYC DYC JLO JMC. Analyzed the data: TYC WLH. Contributed reagents/materials/analysis tools: SSC. Wrote the paper: WLH.