About the Authors

Kazumi Osada

osadak@hoku-iryo-u.ac.jp

Affiliation Division of Physiology, Department of Oral Biology, School of Dentistry, Health Sciences University of Hokkaido, Ishikari-Tobetsu, Hokkaido, Japan

Kenzo Kurihara

Affiliation Aomori University, Kobata, Aomori, Japan

Hiroshi Izumi

Affiliation Division of Physiology, Department of Oral Biology, School of Dentistry, Health Sciences University of Hokkaido, Ishikari-Tobetsu, Hokkaido, Japan

Makoto Kashiwayanagi

Affiliation Department of Sensory Physiology, Asahikawa Medical College, Asahikawa, Hokkaido, Japan

Competing Interests

The authors have declared that no competing interests exist.

Author Contributions

Conceived and designed the experiments: KO KK MK. Performed the experiments: KO MK. Analyzed the data: KO MK. Contributed reagents/materials/analysis tools: KO KK HI MK. Wrote the paper: KO.